REPRODUCTIVE STRATEGIES
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INTRODUCTION
What are the consequences of the major modes of reproduction in trees--
SEXUAL AND ASEXUAL REPRODUCTION
SEXUAL REPRODUCTION
Most widespread form of reproductionTwo step process:
Meiosis-- formation of sex cells (gametes)
cuts number of chromosomes in half important feature is recombination of genes
Fertilization-- union of gametes to form 1st cell of new organism (zygote)
re-establishes "normal" number of chromosomes results in new combinations of genes in same cell due to union of 2 different lines of inheritance (mother, father)
Plants represent nearly a continuum from the union of gametes coming from the same or different individuals.
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Obligate
outcrosser
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outcross/self
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Obligate
selfer
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dioecious
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monoecious,
perfect flowers
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cleistogamous
flowers
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high
heterozygosity
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intermediate
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high
homozygosity
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Examples:
Salix, Populus, Juniperus, Sassafras
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M-
virtually all Gymnosperms, all Amentiferae
PF- most Floriferae |
none
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advantageous
in fluctuating environments; allows production of new genotypes
that might be successful
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advantageous
in stable environmnent; allows close adaptation
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Advantages of sexual reproduction
Variety of progeny in the future Seeds/fruits are formed- allows for effective long-distance dispersal
Disadvantages of sexual reproduction
Too much variation is a problem because it prevents close adaptation to any given environment Seed germination and seedling stages are precarious due to high mortality rates in stressful environment
ASEXUAL REPRODUCTION
Vegetative propagation-- formation of new individual from vegetative parts of old individual
· Stump sprouts-- adventitious buds (buds formed from other meristematic tissue (such as cambium) on stem or branches (when brocken off). e.g Populus, Acer macrophyllum, Quercus, Salix, Castanea dentata.· layering-- roots emerge from branches pressed to round. e.g., Abies lasiocarpa, Thuja plicata, Acer circinatum.
· root crown sprouts-- from top of roots e.g., Sequoia sempervirens.
· root sprouts, suckers-- upright stems emerge from lateral roots. e.g., Populus tremuloides, Populus nigra var. italica, Sassafras albidum.
Advantages of Asexual Reproduction
· well-adapted genotypes can be preserved and multiplied· root sprouts etc take advantage of already existing root system
· no fragile seedling stage \ good for harsh environments
Disadvantages of Asexual Reproduction
· little variation to cope with fluctuating environement (high adaptation, but low adaptibility)· root sprouts-- disease spreads from old to young individuals of clone \ susceptible for fast spread of epidemics
PRACTICAL USE OF ASEXUAL REPRODUCTION
Mass propagation of unique and desireable genotypes. e.g., Populus nigra var. italica- 100% asexual reproduction
Carefully bred genotypes can be propagated e.g. buds of norway spruce are dissected and meristematic tissue grown into seedlings and outplanted
Sterile hybrids can be mass propagated e.g. P.nigra X P. deltoides is desirable tree-- fast growing, but sterile.